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Upstream sequences for all mouse genes were scanned for matches to each of the 1209 motifs, respectively (with masking of protein-coding sequences and repetitive DNA elements)

Multi-omics integration was conducted to reveal core regulatory networks, and mechanistic validation focused on the expression of Nrf2, HO-1, and GPX4

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In addition, the high penetration level through the stratum corneum alone can be explained by the fact that there was no tissue retention from underlying skin that may have limited penetration